DIVISION II. RULES AND RECOMMENDATIONS
CHAPTER III. NOMENCLATURE OF TAXA ACCORDING TO THEIR RANK
SECTION 2. NAMES OF FAMILIES AND SUBFAMILIES, TRIBES AND SUBTRIBES
19.1. The name of a subfamily is a plural adjective used as a noun; it is formed in the same manner as the name of a family
(Art. 18.1) but by using the termination
-oideae instead of -aceae.
19.2. Names intended as names of subfamilies, but published
with their rank denoted by the term "suborder" (subordo) instead of
subfamily, are treated as having been published as names of subfamilies
, unless this would result in a taxonomic sequence with a misplaced rank-denoting term.
Ex. 1. Cyrilloideae
Torr. & A. Gray (Fl. N. Amer. 1: 256. 1838) and
Lindl. (Intr. Nat. Syst. Bot., ed. 2: 238. 1836) were published as
" and "Sub-Order ? Sphenocleaceae
If the term "subfamily" is simultaneously used to denote a rank different
from "suborder", a name published for a taxon at the latter rank cannot be considered to have
been published as the name of a subfamily.
A tribe is designated in a similar manner, with the termination
-eae, and a subtribe similarly with the termination -inae (but not
The name of any subdivision of a family that includes
the type of the adopted, legitimate name of the family to which it is
assigned is to be based on the generic name equivalent to that type
(Art. 10. 6; but see Art. 19.7).
The type of the family name Rosaceae
Juss. is Rosa
L. and hence the subfamily and tribe which include
are to be called Rosoideae
Endl. and Roseae
The type of the family name Gramineae
Juss. (nom. alt.,
Barnhart - see Art. 18.5
) is Poa
L. and hence the subfamily, tribe and subtribe which include
are to be called Pooideae
R. Br. and Poinae
This provision applies only to the names of those
subordinate taxa that include the type of the adopted name of the
family (but see Rec. 19A.2
The subfamily including the type of the family name Ericaceae
L.), irrespective of priority, is to be called
Endl., and the tribe including this type is called Ericeae
D. Don. However, the correct name of the tribe including both
L., the type of the subfamily name Rhododendroideae
L. is Rhodoreae
D. Don (1834) not Rhododendreae
19.5. A name of a subdivision of a family based on an
illegitimate generic name that is not the base of a conserved family
name is illegitimate.
The name Caryophylloideae
Arn. (1832), based on Caryophyllaceae
Juss., nom. cons., is legitimate although it is ultimately based on the illegitimate
Mill. non L.
19.6. When a name of a taxon assigned to one of the above categories has been published with an improper Latin termination, such as
-eae for a subfamily or -oideae for a tribe, the
termination must be changed to accord with the rule, without change of
the author citation or date of publication (see Art. 32.7). However, if such names are published with a non-Latin termination they are not validly published.
Ex. 6. "Climacieae"
(Grout, Moss Fl. N. Amer. 3: 4. 1928), published to designate a subfamily, is to be changed to
However, Melantheen (Kittel in Richard, Nouv. Elém.
Bot., ed. 3, Germ. Transl.: 727. 1840), published to designate a tribe,
is not to be accepted as "Melanthieae
Kitt.", as it has a German rather than a Latin termination. The name
was validly published by Grisebach (Spic. Fl. Rumel. 2: 377. 1846).
19.7. When the Papilionaceae are included in the family Leguminosae (nom. alt.,
Fabaceae; see Art. 18.5) as a subfamily, the name Papilionoideae may be used as an alternative to
19A.1. When a family is changed to the rank of a subdivision
of a family, or the inverse change occurs, and no legitimate name is
available in the new rank, the name should be retained, and only its
(-aceae, -oideae, -eae, -inae) altered.
Ex. 1. The subtribe Drypetinae Griseb. (1859) when raised to the rank of tribe was named
Drypeteae Hurus. (1954); the subtribe Antidesmatinae Müll. Arg. (1865)
when raised to the rank of subfamily was named Antidesmatoideae Hurus. (1954).
19A.2. When a subdivision of a family is changed to another
such rank, and no legitimate name is available in the new rank, its
name should be based on the same generic name as the name in the former
Ex. 2. Among the tribes of the family Ericaceae are
Pyroleae D. Don, Monotropeae D. Don, and Vaccinieae D. Don, none of which includes the type of the family name (Erica L.). The later names
Pyroloideae A. Gray, Monotropoideae A. Gray, and Vaccinioideae Endl. are based on the same generic
(c) 2006, by International Association for Plant Taxonomy. This page last updated